One of five ever built, this is the first of two varients. (the later varient having better engines (Allison 501-D22C turboprops instead of Pratt & Whitney T-34P7 turboprops, for those of you impressed by such stuff)) One remains in service by NASA and the other three are mothballed in England, France, and Germany. (One Guppy per nation) The Super Guppy is a transport vehicle for really big stuff, like plane parts. (In the early 70s, Airbus Industrie used Super Guppies (Boeings at heart) to get their parts to the assembly plants)
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